Creates species combinations to be used in indicator value analyses.
Usage
combinespecies(
X,
min.order = 1,
max.order = 3,
min.occ = 1,
FUN = min,
verbose = FALSE,
add.names = TRUE,
...
)Arguments
- X
A community data table with sites in rows and species in columns. This table can contain either presence-absence or abundance data.
- min.order
Minimum number of species conforming species combinations.
- max.order
Maximum number of species conforming species combinations.
- min.occ
Threshold for minimum occurrence. Combinations with lower values are not kept.
- FUN
Function to be calculated for all species forming the species combination. Accepted values are
min,max,sumandmean.- verbose
If TRUE, prints the results of each step.
- add.names
If TRUE, adds the names of the species combinations to the output matrix. Species combination names are lists of species concatenated using character '+'.
- ...
Additional arguments for function
FUN.
Value
An list with:
XC - A matrix containing the abundance/occurrence of each species combination.
C - A binary matrix describing the set of species forming each combination ('0' means that the species is not included, and '1' means that the species is included).
Details
This function allows creating a data table where rows are sites and columns are combinations of species. Values for a given column of this matrix are derived from the abundance values of the species forming the corresponding combination. In particular, the abundance value for a given combination in a given site is equal to the value returned by function 'FUN' (normally the minimum) among the site values of all species forming the combination. The matrix 'XC' returned by this function can be used in functions strassoc and signassoc. Alternatively, indicators and related functions provide a more elaborated way to explore the indicator value of the simultaneous occurrence of sets of species (i.e. species combinations).
References
De Cáceres, M., Legendre, P., Wiser, S.K. and Brotons, L. 2012. Using species combinations in indicator analyses. Methods in Ecology and Evolution 3(6): 973-982.
Examples
library(stats)
## Loads species data
data(wetland)
## Create species combinations
Y <- combinespecies(X=wetland, max.order=3, min.occ=5, verbose=TRUE)
#> Number of species to combine: 33
#> Starting species 1 ... accepted combinations: 0
#> Starting species 2 ... accepted combinations: 0
#> Starting species 3 ... accepted combinations: 0
#> Starting species 4 ... accepted combinations: 25
#> Starting species 5 ... accepted combinations: 25
#> Starting species 6 ... accepted combinations: 57
#> Starting species 7 ... accepted combinations: 57
#> Starting species 8 ... accepted combinations: 64
#> Starting species 9 ... accepted combinations: 71
#> Starting species 10 ... accepted combinations: 71
#> Starting species 11 ... accepted combinations: 71
#> Starting species 12 ... accepted combinations: 72
#> Starting species 13 ... accepted combinations: 72
#> Starting species 14 ... accepted combinations: 72
#> Starting species 15 ... accepted combinations: 76
#> Starting species 16 ... accepted combinations: 78
#> Starting species 17 ... accepted combinations: 78
#> Starting species 18 ... accepted combinations: 80
#> Starting species 19 ... accepted combinations: 87
#> Starting species 20 ... accepted combinations: 97
#> Starting species 21 ... accepted combinations: 106
#> Starting species 22 ... accepted combinations: 111
#> Starting species 23 ... accepted combinations: 118
#> Starting species 24 ... accepted combinations: 124
#> Starting species 25 ... accepted combinations: 131
#> Starting species 26 ... accepted combinations: 135
#> Starting species 27 ... accepted combinations: 137
#> Starting species 28 ... accepted combinations: 138
#> Starting species 29 ... accepted combinations: 138
#> Starting species 30 ... accepted combinations: 138
#> Starting species 31 ... accepted combinations: 138
#> Starting species 32 ... accepted combinations: 138
#> Starting species 33 ... accepted combinations: 138
#> Final number of combinations: 138
## Creates three site groups using kmeans
wetkm <- kmeans(wetland, centers=3)
## Calculate indicator value of species combinations for each of the three site groups
strassoc(Y$XC, cluster=wetkm$cluster,func="IndVal.g")
#> 1 2 3
#> Pancam 0.27979438 0.07680630 0.78342427
#> Pancam+Melcor 0.29408772 0.09687595 0.73651215
#> Pancam+Melcor+Eupvac 0.13872213 0.11424176 0.77684395
#> Pancam+Melcor+Echpas 0.11451304 0.18860972 0.32063652
#> Pancam+Melcor+Cynarc 0.10403130 0.00000000 0.58257527
#> Pancam+Melcor+Sessp. 0.25354628 0.00000000 0.59160798
#> Pancam+Melcor+Phynod 0.47847438 0.00000000 0.26794565
#> Pancam+Melcor+Echell 0.00000000 0.14969624 0.56904264
#> Pancam+Melcor+Helind 0.42852642 0.08822603 0.14998425
#> Pancam+Melcor+Orysp. 0.32605070 0.17900823 0.15215699
#> Pancam+Eupvac 0.13872213 0.11424176 0.77684395
#> Pancam+Eupvac+Echpas 0.11451304 0.18860972 0.32063652
#> Pancam+Eupvac+Sessp. 0.00000000 0.00000000 0.70710678
#> Pancam+Eupvac+Echell 0.00000000 0.16351749 0.55595945
#> Pancam+Eupvac+Orysp. 0.23792868 0.19594127 0.16655008
#> Pancam+Echpas 0.11451304 0.18860972 0.32063652
#> Pancam+Cynarc 0.08717101 0.00000000 0.59786855
#> Pancam+Sessp. 0.25354628 0.00000000 0.59160798
#> Pancam+Sessp.+Phynod 0.33292257 0.00000000 0.31072773
#> Pancam+Phynod 0.54056248 0.00000000 0.25226249
#> Pancam+Phynod+Helind 0.52822141 0.00000000 0.14790199
#> Pancam+Echell 0.00000000 0.14969624 0.56904264
#> Pancam+Helind 0.49720016 0.08189179 0.13921605
#> Pancam+Helind+Orysp. 0.48675846 0.10021498 0.00000000
#> Pancam+Orysp. 0.36782667 0.15145804 0.18206491
#> Melcor 0.50932091 0.19065489 0.73909205
#> Melcor+Eupvac 0.19306330 0.10599553 0.81086584
#> Melcor+Eupvac+Echpas 0.19081501 0.15714177 0.40071151
#> Melcor+Eupvac+Cynarc 0.00000000 0.00000000 0.70710678
#> Melcor+Eupvac+Sessp. 0.00000000 0.00000000 0.77459667
#> Melcor+Eupvac+Phynod 0.33292257 0.00000000 0.31072773
#> Melcor+Eupvac+Echell 0.00000000 0.16351749 0.55595945
#> Melcor+Eupvac+Orysp. 0.32605070 0.17900823 0.15215699
#> Melcor+Echpas 0.25047110 0.13751354 0.46754605
#> Melcor+Aesind 0.46004371 0.00000000 0.16101530
#> Melcor+Aesind+Phynod 0.46004371 0.00000000 0.16101530
#> Melcor+Aesind+Helind 0.46004371 0.00000000 0.16101530
#> Melcor+Cyprot 0.21466940 0.00000000 0.45080573
#> Melcor+Cyprot+Sessp. 0.21466940 0.00000000 0.45080573
#> Melcor+Cynarc 0.08132501 0.00000000 0.73786479
#> Melcor+Cynarc+Sessp. 0.08717101 0.00000000 0.66843736
#> Melcor+Sessp. 0.32387780 0.05334458 0.67863027
#> Melcor+Sessp.+Phynod 0.47847438 0.00000000 0.26794565
#> Melcor+Sessp.+Helind 0.46004371 0.00000000 0.16101530
#> Melcor+Phynod 0.79138739 0.00000000 0.20144406
#> Melcor+Phynod+Echell 0.46004371 0.00000000 0.16101530
#> Melcor+Phynod+Helind 0.74586774 0.00000000 0.11602387
#> Melcor+Phynod+Elesp. 0.70710678 0.00000000 0.00000000
#> Melcor+Echell 0.28523530 0.11744983 0.48907666
#> Melcor+Echell+Helind 0.42852642 0.08822603 0.14998425
#> Melcor+Helind 0.69300757 0.12682491 0.10780118
#> Melcor+Helind+Ipoaqu 0.35081263 0.09630150 0.16371256
#> Melcor+Helind+Orysp. 0.37193595 0.20420013 0.00000000
#> Melcor+Helind+Elesp. 0.61387889 0.08425789 0.00000000
#> Melcor+Ipoaqu 0.29380883 0.16130681 0.27422158
#> Melcor+Orysp. 0.34652229 0.35671412 0.12128280
#> Melcor+Elesp. 0.63620901 0.14969624 0.00000000
#> Eupvac 0.24480142 0.12345500 0.77112447
#> Eupvac+Echpas 0.26956145 0.14799452 0.37738603
#> Eupvac+Cynarc 0.00000000 0.00000000 0.70710678
#> Eupvac+Sessp. 0.08717101 0.00000000 0.73223645
#> Eupvac+Phynod 0.40996003 0.00000000 0.28697202
#> Eupvac+Echell 0.00000000 0.16351749 0.55595945
#> Eupvac+Orysp. 0.30588648 0.20568080 0.14274702
#> Echpas 0.35850111 0.23618896 0.40152124
#> Echpas+Sessp. 0.11451304 0.18860972 0.32063652
#> Echpas+Phynod 0.59761430 0.00000000 0.00000000
#> Echpas+Echell 0.25280383 0.31228708 0.00000000
#> Echpas+Helind 0.34279953 0.28230550 0.00000000
#> Echpas+Helind+Orysp. 0.25280383 0.31228708 0.00000000
#> Echpas+Orysp. 0.31958752 0.35091963 0.00000000
#> Carhal 0.38959686 0.18523964 0.00000000
#> Aesind 0.53812200 0.00000000 0.13754606
#> Aesind+Phynod 0.53812200 0.00000000 0.13754606
#> Aesind+Phynod+Helind 0.52822141 0.00000000 0.14790199
#> Aesind+Helind 0.52822141 0.00000000 0.14790199
#> Cyprot 0.26291524 0.00000000 0.45080573
#> Cyprot+Sessp. 0.29880715 0.00000000 0.41833001
#> Cynarc 0.04739390 0.00000000 0.76232019
#> Cynarc+Sessp. 0.08717101 0.00000000 0.66843736
#> Walind 0.21438482 0.30579739 0.32878553
#> Walind+Sessp. 0.16640855 0.34260584 0.34945796
#> Walind+Sessp.+Orysp. 0.00000000 0.54232614 0.00000000
#> Walind+Sessp.+Ludads 0.00000000 0.54232614 0.00000000
#> Walind+Orysp. 0.00000000 0.54232614 0.00000000
#> Walind+Orysp.+Ludads 0.00000000 0.54232614 0.00000000
#> Walind+Ludads 0.00000000 0.54232614 0.00000000
#> Sessp. 0.32684653 0.38066066 0.52837381
#> Sessp.+Phynod 0.55328334 0.00000000 0.23904572
#> Sessp.+Phynod+Helind 0.46004371 0.00000000 0.16101530
#> Sessp.+Echell 0.12429372 0.14475824 0.42623865
#> Sessp.+Helind 0.36256278 0.25857883 0.12689697
#> Sessp.+Ipoaqu 0.11451304 0.18860972 0.32063652
#> Sessp.+Orysp. 0.08101671 0.61149560 0.00000000
#> Sessp.+Orysp.+Ludads 0.00000000 0.64168895 0.00000000
#> Sessp.+Elesp. 0.31958752 0.30390531 0.00000000
#> Sessp.+Ludads 0.00000000 0.64168895 0.00000000
#> Phynod 0.96324195 0.00000000 0.12013738
#> Phynod+Echell 0.55593699 0.00000000 0.11602387
#> Phynod+Echell+Helind 0.52822141 0.00000000 0.14790199
#> Phynod+Echell+Elesp. 0.59761430 0.00000000 0.00000000
#> Phynod+Helind 0.84221458 0.00000000 0.09860133
#> Phynod+Helind+Orysp. 0.59761430 0.00000000 0.00000000
#> Phynod+Helind+Elesp. 0.75592895 0.00000000 0.00000000
#> Phynod+Orysp. 0.65465367 0.00000000 0.00000000
#> Phynod+Elesp. 0.80178373 0.00000000 0.00000000
#> Echell 0.39411120 0.19353149 0.40294538
#> Echell+Helind 0.47107212 0.15517670 0.13190019
#> Echell+Helind+Elesp. 0.59761430 0.00000000 0.00000000
#> Echell+Orysp. 0.11200122 0.36894520 0.15680171
#> Echell+Elesp. 0.55374887 0.09120570 0.00000000
#> Helind 0.67175503 0.41377764 0.07864497
#> Helind+Ipoaqu 0.34875678 0.16582162 0.14094837
#> Helind+Orysp. 0.40508353 0.50435307 0.00000000
#> Helind+Orysp.+Elesp. 0.32494624 0.34547372 0.00000000
#> Helind+Orysp.+Ludads 0.00000000 0.59408853 0.00000000
#> Helind+Elesp. 0.62614405 0.27177038 0.00000000
#> Helind+Ludads 0.00000000 0.59408853 0.00000000
#> Ipoaqu 0.28019610 0.33305854 0.22647981
#> Ipoaqu+Orysp. 0.15281246 0.44493123 0.00000000
#> Ipoaqu+Orysp.+Ludads 0.00000000 0.54232614 0.00000000
#> Ipoaqu+Elesp. 0.38959686 0.18523964 0.00000000
#> Ipoaqu+Psespi 0.25280383 0.31228708 0.00000000
#> Ipoaqu+Ludads 0.00000000 0.54232614 0.00000000
#> Orysp. 0.30880860 0.85614502 0.06671028
#> Orysp.+Elesp. 0.30159199 0.56637046 0.00000000
#> Orysp.+Elesp.+Psespi 0.00000000 0.54232614 0.00000000
#> Orysp.+Elesp.+Ludads 0.00000000 0.64168895 0.00000000
#> Orysp.+Psespi 0.00000000 0.64168895 0.00000000
#> Orysp.+Psespi+Ludads 0.00000000 0.64168895 0.00000000
#> Orysp.+Ludads 0.00000000 0.90748521 0.00000000
#> Elesp. 0.54643833 0.50200498 0.00000000
#> Elesp.+Psespi 0.18239545 0.47499947 0.00000000
#> Elesp.+Psespi+Ludads 0.00000000 0.54232614 0.00000000
#> Elesp.+Ludads 0.00000000 0.64168895 0.00000000
#> Psespi 0.13031890 0.60234015 0.00000000
#> Psespi+Ludads 0.00000000 0.64168895 0.00000000
#> Ludads 0.00000000 0.90748521 0.00000000
## Calculate point biserial correlation value of species combinations
## for each of the three site groups
strassoc(Y$XC, cluster=wetkm$cluster,func="r.g")
#> 1 2 3
#> Pancam -0.187860796 -0.353094935 0.540955730
#> Pancam+Melcor -0.153571185 -0.427119859 0.580691045
#> Pancam+Melcor+Eupvac -0.311569734 -0.348858665 0.660428398
#> Pancam+Melcor+Echpas -0.122627314 -0.025350839 0.147978153
#> Pancam+Melcor+Cynarc -0.166538214 -0.305320060 0.471858274
#> Pancam+Melcor+Sessp. -0.039528471 -0.395284708 0.434813178
#> Pancam+Melcor+Phynod 0.311714600 -0.337690817 0.025976217
#> Pancam+Melcor+Echell -0.303450465 -0.130050199 0.433500665
#> Pancam+Melcor+Helind 0.273735074 -0.177844173 -0.095890902
#> Pancam+Melcor+Orysp. 0.141612141 -0.053034029 -0.088578112
#> Pancam+Eupvac -0.311569734 -0.348858665 0.660428398
#> Pancam+Eupvac+Echpas -0.122627314 -0.025350839 0.147978153
#> Pancam+Eupvac+Sessp. -0.316227766 -0.316227766 0.632455532
#> Pancam+Eupvac+Echell -0.322901537 -0.102741398 0.425642936
#> Pancam+Eupvac+Orysp. 0.049340939 -0.005482327 -0.043858612
#> Pancam+Echpas -0.122627314 -0.025350839 0.147978153
#> Pancam+Cynarc -0.176255712 -0.258875577 0.435131290
#> Pancam+Sessp. -0.039528471 -0.395284708 0.434813178
#> Pancam+Sessp.+Phynod 0.156158885 -0.283037978 0.126879094
#> Pancam+Phynod 0.380594451 -0.364046866 -0.016547585
#> Pancam+Phynod+Helind 0.402873283 -0.299812676 -0.103060607
#> Pancam+Echell -0.303450465 -0.130050199 0.433500665
#> Pancam+Helind 0.346936555 -0.212045943 -0.134890612
#> Pancam+Helind+Orysp. 0.378947780 -0.124245174 -0.254702606
#> Pancam+Orysp. 0.125455012 -0.123793356 -0.001661656
#> Melcor -0.008864143 -0.585033456 0.593897600
#> Melcor+Eupvac -0.282236493 -0.421140331 0.703376824
#> Melcor+Eupvac+Echpas -0.079780769 -0.125514968 0.205295738
#> Melcor+Eupvac+Cynarc -0.316227766 -0.316227766 0.632455532
#> Melcor+Eupvac+Sessp. -0.353553391 -0.353553391 0.707106781
#> Melcor+Eupvac+Phynod 0.156158885 -0.283037978 0.126879094
#> Melcor+Eupvac+Echell -0.322901537 -0.102741398 0.425642936
#> Melcor+Eupvac+Orysp. 0.141612141 -0.053034029 -0.088578112
#> Melcor+Echpas -0.048885532 -0.207994104 0.256879636
#> Melcor+Aesind 0.331964402 -0.271607238 -0.060357164
#> Melcor+Aesind+Phynod 0.331964402 -0.271607238 -0.060357164
#> Melcor+Aesind+Helind 0.331964402 -0.271607238 -0.060357164
#> Melcor+Cyprot -0.009492435 -0.294265490 0.303757925
#> Melcor+Cyprot+Sessp. -0.009492435 -0.294265490 0.303757925
#> Melcor+Cynarc -0.258692595 -0.358189746 0.616882341
#> Melcor+Cynarc+Sessp. -0.219766536 -0.322782100 0.542548636
#> Melcor+Sessp. -0.061415718 -0.441693273 0.503108990
#> Melcor+Sessp.+Phynod 0.311714600 -0.337690817 0.025976217
#> Melcor+Sessp.+Helind 0.331964402 -0.271607238 -0.060357164
#> Melcor+Phynod 0.688211830 -0.494652253 -0.193559577
#> Melcor+Phynod+Echell 0.331964402 -0.271607238 -0.060357164
#> Melcor+Phynod+Helind 0.647489722 -0.405656212 -0.241833511
#> Melcor+Phynod+Elesp. 0.632455532 -0.316227766 -0.316227766
#> Melcor+Echell -0.057767702 -0.256966673 0.314734375
#> Melcor+Echell+Helind 0.273735074 -0.177844173 -0.095890902
#> Melcor+Helind 0.556610474 -0.264510406 -0.292100068
#> Melcor+Helind+Ipoaqu 0.192665548 -0.140447783 -0.052217765
#> Melcor+Helind+Orysp. 0.233623386 0.015785364 -0.249408750
#> Melcor+Helind+Elesp. 0.510104338 -0.198672216 -0.311432122
#> Melcor+Ipoaqu 0.068454989 -0.110461460 0.042006471
#> Melcor+Orysp. 0.099829752 0.114023556 -0.213853308
#> Melcor+Elesp. 0.514344500 -0.154303350 -0.360041150
#> Eupvac -0.222162428 -0.366327731 0.588490160
#> Eupvac+Echpas 0.006314645 -0.161266318 0.154951673
#> Eupvac+Cynarc -0.316227766 -0.316227766 0.632455532
#> Eupvac+Sessp. -0.258518870 -0.379699591 0.638218461
#> Eupvac+Phynod 0.237663538 -0.310790780 0.073127242
#> Eupvac+Echell -0.322901537 -0.102741398 0.425642936
#> Eupvac+Orysp. 0.085445371 0.021715398 -0.107160769
#> Echpas 0.039571256 -0.143551045 0.103979789
#> Echpas+Sessp. -0.122627314 -0.025350839 0.147978153
#> Echpas+Phynod 0.519875245 -0.259937622 -0.259937622
#> Echpas+Echell 0.083491298 0.160560189 -0.244051487
#> Echpas+Helind 0.176542380 0.097098309 -0.273640690
#> Echpas+Helind+Orysp. 0.083491298 0.160560189 -0.244051487
#> Echpas+Orysp. 0.127631126 0.169249971 -0.296881097
#> Carhal 0.256673784 -0.028519309 -0.228154475
#> Aesind 0.406467535 -0.283760355 -0.122707180
#> Aesind+Phynod 0.406467535 -0.283760355 -0.122707180
#> Aesind+Phynod+Helind 0.402873283 -0.299812676 -0.103060607
#> Aesind+Helind 0.402873283 -0.299812676 -0.103060607
#> Cyprot -0.009224789 -0.285968450 0.295193238
#> Cyprot+Sessp. 0.080408440 -0.321633760 0.241225320
#> Cynarc -0.328105913 -0.362283613 0.690389526
#> Cynarc+Sessp. -0.219766536 -0.322782100 0.542548636
#> Walind -0.011527563 -0.015286551 0.026814114
#> Walind+Sessp. -0.168862453 0.079600012 0.089262441
#> Walind+Sessp.+Orysp. -0.233126202 0.466252404 -0.233126202
#> Walind+Sessp.+Ludads -0.233126202 0.466252404 -0.233126202
#> Walind+Orysp. -0.219970673 0.439941345 -0.219970673
#> Walind+Orysp.+Ludads -0.233126202 0.466252404 -0.233126202
#> Walind+Ludads -0.233126202 0.466252404 -0.233126202
#> Sessp. -0.152623053 0.033719047 0.118904007
#> Sessp.+Phynod 0.396197766 -0.346673045 -0.049524721
#> Sessp.+Phynod+Helind 0.331964402 -0.271607238 -0.060357164
#> Sessp.+Echell -0.102225251 -0.135559572 0.237784823
#> Sessp.+Helind 0.123418836 0.044360507 -0.167779343
#> Sessp.+Ipoaqu -0.122627314 -0.025350839 0.147978153
#> Sessp.+Orysp. -0.194301278 0.462553042 -0.268251764
#> Sessp.+Orysp.+Ludads -0.282038037 0.564076075 -0.282038037
#> Sessp.+Elesp. 0.111605128 0.147998104 -0.259603232
#> Sessp.+Ludads -0.282038037 0.564076075 -0.282038037
#> Phynod 0.936179717 -0.524910015 -0.411269702
#> Phynod+Echell 0.428999293 -0.268770641 -0.160228652
#> Phynod+Echell+Helind 0.402873283 -0.299812676 -0.103060607
#> Phynod+Echell+Elesp. 0.519875245 -0.259937622 -0.259937622
#> Phynod+Helind 0.731521080 -0.428207461 -0.303313618
#> Phynod+Helind+Orysp. 0.487122300 -0.243561150 -0.243561150
#> Phynod+Helind+Elesp. 0.652071200 -0.326035600 -0.326035600
#> Phynod+Orysp. 0.527002968 -0.263501484 -0.263501484
#> Phynod+Elesp. 0.703597545 -0.351798772 -0.351798772
#> Echell 0.134680432 -0.230916637 0.096236205
#> Echell+Helind 0.297437874 -0.132865649 -0.164572225
#> Echell+Helind+Elesp. 0.519875245 -0.259937622 -0.259937622
#> Echell+Orysp. -0.132495108 0.205974927 -0.073479820
#> Echell+Elesp. 0.447048298 -0.163344570 -0.283703727
#> Helind 0.424041854 0.053653031 -0.477694885
#> Helind+Ipoaqu 0.192700899 -0.081932211 -0.110768688
#> Helind+Orysp. 0.152537359 0.250597089 -0.403134448
#> Helind+Orysp.+Elesp. 0.132661776 0.144721937 -0.277383713
#> Helind+Orysp.+Ludads -0.258198890 0.516397779 -0.258198890
#> Helind+Elesp. 0.425572844 -0.023442572 -0.402130272
#> Helind+Ludads -0.258198890 0.516397779 -0.258198890
#> Ipoaqu 0.037196844 0.049326249 -0.086523093
#> Ipoaqu+Orysp. -0.004775024 0.253076289 -0.248301265
#> Ipoaqu+Orysp.+Ludads -0.233126202 0.466252404 -0.233126202
#> Ipoaqu+Elesp. 0.264116315 -0.029346257 -0.234770058
#> Ipoaqu+Psespi 0.083491298 0.160560189 -0.244051487
#> Ipoaqu+Ludads -0.233126202 0.466252404 -0.233126202
#> Orysp. -0.203376016 0.733434767 -0.530058751
#> Orysp.+Elesp. -0.016527703 0.384269088 -0.367741386
#> Orysp.+Elesp.+Psespi -0.218750000 0.437500000 -0.218750000
#> Orysp.+Elesp.+Ludads -0.282038037 0.564076075 -0.282038037
#> Orysp.+Psespi -0.258198890 0.516397779 -0.258198890
#> Orysp.+Psespi+Ludads -0.282038037 0.564076075 -0.282038037
#> Orysp.+Ludads -0.418609179 0.837218358 -0.418609179
#> Elesp. 0.188565688 0.290705436 -0.479271124
#> Elesp.+Psespi -0.082302966 0.355399171 -0.273096205
#> Elesp.+Psespi+Ludads -0.219970673 0.439941345 -0.219970673
#> Elesp.+Ludads -0.267859812 0.535719624 -0.267859812
#> Psespi -0.176687998 0.451322603 -0.274634605
#> Psespi+Ludads -0.267859812 0.535719624 -0.267859812
#> Ludads -0.351927849 0.703855698 -0.351927849